A beautiful market town in the heart of Lincolnshire’s dramatic Fenland, Boston is immersed in history and was the setting for a dramatic twist in the Pilgrims' history.
Leiden American Pilgrim Museum
After escaping England, the Dutch city of Leiden offered sanctuary to the Pilgrims and the promise of a new life. Leiden is as welcoming now as it was then.
Scrooby Manor House
The birthplace of Pilgrim elder William Brewster, Scrooby & Babworth is a firm part of the origins of the Mayflower story. These pretty villages are in the heart of rural Nottinghamshire.
St. Helena's Church, Austerfield
The second elected Governor of the Plymouth Colony in Massachusetts was William Bradford, from Austerfield - a small village near Doncaster that boasts superb English countryside.
Dartmouth is a renowned coastal destination and a hub of picturesque places to eat and drink. The Speedwell stopped here for repairs before travelling to Plymouth.
The Great Hall, Gainsborough Old Hall
Some of the Separatists are thought to have worshipped clandestinely at Gainsborough Old Hall - now regarded as one of the best preserved medieval manor houses in Britain.
Low Lighthouse, Harwich. Essex.
This attractive maritime town is the home of the Mayflower, where the historic ship was originally built. It is where its captain, Christopher Jones lived and was twice wed
Ross Castle, Cleethorpes
The Separatists escaped to Holland from Immingham Creek, now a busy port near the popular town of Cleethorpes that's bursting with history.
Smeatons Tower, Plymouth Hoe
The final stop before the Pilgrims crossed the Atlantic, Britain's Ocean City is one of England's fastest growing destinations, home to the famous Mayflower Steps.
Mayflower Pub, Rotherhithe
The home of the Mayflower's captain and much of its crew, Southwark is an internationally-recognised cultural destination, known for its vibrant performing arts.
Mayflower Memorial, Southampton
The port where the Mayflower and Speedwell met, Southampton is steeped in maritime heritage and today is known as the cruise capital of Europe.
St. Peters Church, Droitwich
The birthplace of Edward Winslow, a senior leader on the Mayflower, Worcestershire is home to an amazing evolving history, including the first and last battle of the Civil War.
Mullins family plaque
Dorking is a quintessentially English town in Surrey, home to six Mayflower passengers including William Mullins - his house still stands in the town today
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Austerfield, near Doncaster and the market town of Bawtry. Home of William Bradford who was baptised in St. Helena's Church and later became Plymouth Colony's first elected Govenor.
In the Bassetlaw area of north Nottinghamshire, the beliefs of the leading Separatists who voyaged to American in 1620 were shaped. Explore the Pilgrim Roots.
Gainsborough, Lincolnshire. Close to Scrooby and Retford. Some of the Separatists are thought to have worshipped in secret at the Old Hall with permission of its owner, merchant William Hickman.
The Scrooby congregation - including Williams Brewster and Bradford - made their first attempt to escape to Holland from Boston in Lincolnshire. They were held and tried in Boston Guildhall.
It was from Immingham Creek on the edge of the river Humber that the Separatists made a dangerous, but successful escape from England to Holland in their search for religious freedom.
Prominent Mayflower passenger, Edward Winslow came from Worcestershire - he was schooled in Worcester and came from Droitwich Spa. Following the voyage, he later returned to England.
The Separatists escaped from England to Holland and settled in Leiden - a city of free-thinkers and religious tolerance. It was their resting place for almost 12 years before departing on the Speedwell in preparation for their final voyage to the New World.
The Mayflower is believed to have been built in Harwich sometime before 1600, and was commanded and part-owned by her Master, Captain Christopher Jones, whose house still stands near the waterfront.
The London borough of Southwark, which includes the former docklands of Rotherhithe, has many links with the voyage of the Pilgrims. It was the home port of the Mayflower, and the area was one with its own strong tradition of religious descent.
The Mayflower arrived in Southampton in July 1620 and several days later was joined by the Speedwell, carrying the Pilgrims from Leiden. On 15 August the two ships weighted anchor and set sail.
When the Speedwell began to take on water, both ships and their crew changed course and arrived in Dartmouth on 23 August. They rested here whilst repairs were made in Bayards cove before heading out into the English Channel.
300 miles clear of England, the Speedwell continued to leak and both ships turned about for Plymouth. Eventually, just the Mayflower set sail with up to 30 crew and 102 passengers on board. The final departure city before embarking on life in the New World.
After a storm tossed 66 days at sea, the Mayflower anchored on the tip of Cape Cod, at what is now Provincetown, Massachusetts. The settlers formulated the 'Mayflower Compact'.